Greg Copenhaver

• Assistant Professor, The University of North Carolina, Chapel Hill, North Carolina, Dept. of Biology
• Postdoctoral research in the lab of Dr. Daphne Preuss, University of Chicago, Illinois, present position.
• Ph.D. in biology and biomedical sciences, Washington University St. Louis, Missouri, August, 1996.
• B.S. in Botany with high distinction, University of California Riverside, California, 1990.

• American Association for the Advancement of Science
• American Society for Cell Biology
• Genetics Society of America
• Sigma Xi

• Academic Program Service Award, U.C. Riverside, 1989.
• Bernarr J. Hall Scholarship, 1989-90.
• James and Adelaine Wallace Annual Prize for Botany, 1990.
• Botanical Society of America Young Botanist Award, 1990.
• U.C. Riverside nominee for Rhodes-Marshall Scholarship, 1990.
• Outreach Service Recognition Award-Beaumont High School, 1994
• Trainee on NSF training grant, 1994-95.
• Monsanto Predoctoral Fellowship, 1995-96.
• USDA grant (2 years, $82,000) 1997-98

• Teaching Assistant, Washington University, Biology 337, fall 1992.
• Outreach Teaching Assistant, Beaumont High School, St. Louis, Missouri, spring 1994.
• Graduate Student Mentor for Summer Scholars in Biology and Biomedical Sciences Program, Washington University, summer 1994 and 1995.
• Instructor - Genetic Analysis, University of Chicago, fall 1998

• Copenhaver, G.P., Putnam, C.D., Denton, M., Pikaard, C.S. (1994) The RNA polymerase I transcription factor UBF is a sequence-tolerant HMG-box protein that can recognize structured nucleic acids. Nucleic Acids Research 22(13): 2651-2657.


• Putnam, C.D., Copenhaver, G.P., Denton M., Pikaard, C.S. (1994 ) The RNA polymerase I transactivator upstream binding factor requires its dimerization domain and high-mobility-group (HMG) box 1 to bend, wrap and positively supercoil enhancer DNA. Molecular and Cellular Biology 14(10): 6476-6488.


• Copenhaver, G.P., Doelling, J.H., Gens, S.J., Pikaard, C.S. (1995) Use of RFLPs larger than 100 kbp to map the position and internal organization of the nucleolus organizer region on chromosome 2 in Arabidopsis thaliana. The Plant Journal 7(2): 273-286.


• Copenhaver, G.P., and Pikaard, C.S. (1996) RFLP and physical mapping with an rDNA-specific endonuclease reveals that nucleolus organizer regions of Arabidopsis thaliana adjoin the telomeres on chromosomes 2 and 4.The Plant Journal 9(2): 259-272.


• Copenhaver, G.P. and Pikaard, C.S. (1996) Two-dimensional RFLP analyses reveal megabase-sized clusters of rRNA gene variants in Arabidopsis thaliana, suggesting local spreading of variants as the mode for gene homogenization during concerted evolution. The Plant Journal 9(2): 273-282.


• Copenhaver, G.P., Browne, W.E. and Preuss, D. (1998 ) Assaying genome-wide recombination and centromere functions with Arabidopsis tetrads. Proc. Natl. Acad. Sci 95:247-252.


• Copenhaver, G.P. and Preuss, D. (1999) Centromeres in the genomic era: Unraveling paradoxes. Curr. Op. Plant Biol. 2: 104-108.

• Plant centromere compositions and methods, G.P. Copenhaver and D. Preuss, University of Chicago, (pending, 1999).


• Chemical and physical treatmenst that stimuate recombination, G.P. Copenhaver and D. Preuss, The University of Chicago, (pending, 1999)

• Copenhaver, G.P., Browne, W.E. and Preuss, D. Mapping Centromeres in Arabidopsis using genome-wide recombination analysis. 1997 8th International Conference on Arabidopsis Research, Madison Wisconsin.


• Copenhaver, G.P., Nickel, K., and Preuss, D. Assaying Centromere functions and genome-wide recombination in Arabidopsis. 1998 Plant and Animal Genome VI conference, San Diego, California.


• Copenhaver, G.P., Kuramori, T., Nickel, K., Gordon, I. and Daphne Preuss. Genetic and Physical Characterization of Arabidopsis Centromeres. 1998 9th International Conference on Arabidopsis Research, Madison Wisconsin.


• Copenhaver, G.P., Kuramori, T., Nickel, K. and Daphne Preuss. The Five Arabidopsis Centromeres: Genetic Location and Sequence Composition. 1998 Biotechnology Conference at Cold Spring Harbor, Cold Spring Harbor, New York.

• Putnam, C.D., Copenhaver, G.P., Pikaard, C.S. Minimum DNA sequence requirements for xUBF binding in vitro. 1991 Synthesis of Ribosomes meeting, Cold Spring Harbor.


• Putnam, C.D., Copenhaver, G.P., Denton, M.L., Pikaard, C.S. The RNA polymerase I trans-activator UBF requires the dimerization domain and HMG-box 1 to bend, wrap, and positively supercoil enhancer DNA. 1994 Ribosome Synthesis and Nucleolar Function meeting, Cold Spring Harbor.


• Copenhaver, G.P., Doelling, J.H., Pikaard, C.S. rRNA gene variants are both clustered and dispersed on chromosome 2 in A. thaliana - RFLP analysis on pulsed-field gels as a method for mapping co-evolving repetitive gene clusters with limited sequence heterogeneity. 1994 Ribosome Synthesis and Nucleolar Function meeting, Cold Spring Harbor.


• Copenhaver, G.P., Pikaard, C.S. Organization and Co-evolution of 6 Mbp of the Arabidopsis genome: RFLP and physical mapping of the nucleolus organizer regions and associated telomeres on chromosomes 2 and 4 using 1- and 2-D pulsed-field gels. 1995 6th International Conference on Arabidopsis Research, Madison Wisconsin.


• Copenhaver, G.P., Pikaard, C.S. RFLP and physical mapping with an rDNA specific endonuclease reveals that nucleolus organizer regions of Arabidopsis thaliana adjoin the telomeres on chromosomes 2 and 4. 1996 International Plant Genome Conference - Plant Genome IV, San Diego, California.


• Copenhaver, G.P., Pikaard, C.S. Distribution of four classes of rRNA gene variants within the multi-megabase nucleolus organizer regions of Arabidopsis thaliana chromosomes 2 and 4. 1996 International Plant Genome Conference - Plant Genome IV, San Diego, California


• Copenhaver, G.P., Browne, W.E. and Preuss, D. Measuring genome-wide recombination and centromere functions in Arabidopsis meiosis. 1997 37th American Society for Cell Biology Annual Meeting , Washington D.C.